Clarks Size Calculator

Population Size Calculator

Enter how many animals you caught and marked the first time, how many you caught the second time, and how many of those already had a mark. The calculator estimates the population and gives a 95% range.

  • Mark and recapture
  • 95% range
  • Chapman's method
Your survey

From 1 to 10000000

From 1 to 10000000

From 0 to 10000000

Leave enough time between catches for marked animals to mix back in, but not so long that many are born or die.

You can count a population without catching every animal. Mark a sample, release them and catch again later. If 10 of your second catch of 100 are marked, the 100 marked animals are about a tenth of the population, so there are about 1,000.

Population estimate chart

Chapman's estimator. Search for a survey.

Estimates
MarkedSecond catchRecapturedEstimate95% range
50505433149 to 716
10010010926459 to 1,394
10010020485324 to 646
10010050199172 to 226
20015030978700 to 1,256

The range narrows as recaptures increase. Aim for at least 7, ideally 20 or more.

Recaptures and estimate

100 marked, 100 caught. Slide the recaptures.

The dots are the second catch. Coloured dots are animals that already carried a mark.

How mark and recapture works

Mark, release, recatch. Step through it.

First catch

Count and mark.

Use marks that do not harm the animal or change its chance of being caught, such as a small paint spot or tag.

Lincoln-Petersen, Chapman or Schnabel

Methods. Compare them.

M × C ÷ R.

CatchesTwo
BiasHigh with few recaptures
UseTeaching

The Schnabel method uses several catches in a row and gives a tighter estimate for the same effort.

Before you survey

Field checks. Tick each item.

Tick each item as you go.

If marked animals avoid traps after the first catch, the estimate will be too high.

What you get

Population estimate. Each tile explains one result.

Estimate

Chapman.

95% range

Low to high.

Simple estimate

Lincoln-Petersen.

Marked share

Of second catch.

Precision

Low to good.

Small recapture numbers give wide ranges. Doubling the effort in the second catch helps more than anything else.

Frequently Asked Questions

How does mark and recapture work?

Mark a sample, release it, recatch, and use the share of marked animals to estimate the total.

What is the Lincoln-Petersen formula?

Population = marked × second catch ÷ recaptured.

Why use Chapman's estimator?

It is less biased when recaptures are few, and works even with none.

How many recaptures do I need?

At least 7 for a rough figure, 20 or more for a good one.

What if no marked animals are recaptured?

The population is probably large. Catch more animals next time.

Does mark and recapture work for fish?

Yes, it is widely used for fish in ponds and lakes.

What assumptions does it make?

No births, deaths or moves, no lost marks and equal catchability.

Count what you cannot see

Two catches and a careful count give a sound estimate of the whole population. More recaptures give a tighter answer.

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